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@PHDTHESIS{vanMeegen:1010679,
author = {van Meegen, Alexander},
title = {{S}imulation and theory of large-scale cortical networks},
volume = {98},
school = {Univ. Köln},
type = {Dissertation},
address = {Jülich},
publisher = {Forschungszentrum Jülich GmbH Zentralbibliothek, Verlag},
reportid = {FZJ-2023-03187},
isbn = {978-3-95806-708-0},
series = {Schriften des Forschungszentrums Jülich Reihe Information
/ Information},
pages = {250 p.},
year = {2023},
note = {Dissertation, Univ. Köln, 2022},
abstract = {Cerebral cortex is composed of intricate networks of
neurons. These neuronal networks are strongly
interconnected: every neuron receives, on average, input
from thousands or more presynaptic neurons. In fact, to
support such a number of connections, a majority of the
volume inthe cortical gray matter is filled by axons and
dendrites. Besides the networks, neurons themselves are also
highly complex. They possess an elaborate spatial structure
and support various types of active processes and
nonlinearities. In the face of such complexity, it seems
necessary to abstract away some of the details and to
investigate simplified models.In this thesis, such
simplified models of neuronal networks are examined on
varying levels of abstraction. Neurons are modeled as point
neurons, both rate-based and spike-based, and networks are
modeled as block-structured random networks. Crucially, on
this level of abstraction, the models are still amenable to
analytical treatment using the framework of dynamical
mean-field theory.The main focus of this thesis is to
leverage the analytical tractability of random networks of
point neurons in order to relate the network structure, and
the neuron parameters, to the dynamics of the neurons—in
physics parlance, to bridge across the scales from neurons
to networks.More concretely, four different models are
investigated: 1) fully connected feedforward networks and
vanilla recurrent networks of rate neurons; 2)
block-structured networks of rate neurons in continuous
time; 3) block-structured networks of spiking neurons; and
4) a multi-scale, data-based network of spiking neurons. We
consider the first class of models in the light of Bayesian
supervised learning and compute their kernel in the
infinite-size limit. In the second class of models, we
connect dynamical mean-field theory with large-deviation
theory, calculate beyond mean-field fluctuations, and
perform parameter inference. For the third class of models,
we develop a theory for the autocorrelation time of the
neurons. Lastly, we consolidate data across multiple
modalities into a layer- and population-resolved model of
human cortex and compare its activity with cortical
recordings.In two detours from the investigation of these
four network models, we examine the distribution of neuron
densities in cerebral cortex and present a software toolbox
for mean-field analyses of spiking networks.},
cin = {INM-6 / IAS-6 / INM-10},
cid = {I:(DE-Juel1)INM-6-20090406 / I:(DE-Juel1)IAS-6-20130828 /
I:(DE-Juel1)INM-10-20170113},
pnm = {5231 - Neuroscientific Foundations (POF4-523) / 5232 -
Computational Principles (POF4-523) / DFG project 313856816
- SPP 2041: Computational Connectomics (313856816) / DFG
project 347572269 - Heterogenität von Zytoarchitektur,
Chemoarchitektur und Konnektivität in einem großskaligen
Computermodell der menschlichen Großhirnrinde (347572269) /
Brain-Scale Simulations $(jinb33_20090701)$ / Brain-Scale
Simulations $(jinb33_20191101)$ / Brain-Scale Simulations
$(jinb33_20220812)$ / HBP SGA2 - Human Brain Project
Specific Grant Agreement 2 (785907) / HBP SGA3 - Human Brain
Project Specific Grant Agreement 3 (945539)},
pid = {G:(DE-HGF)POF4-5231 / G:(DE-HGF)POF4-5232 /
G:(GEPRIS)313856816 / G:(GEPRIS)347572269 /
$G:(DE-Juel1)jinb33_20090701$ /
$G:(DE-Juel1)jinb33_20191101$ /
$G:(DE-Juel1)jinb33_20220812$ / G:(EU-Grant)785907 /
G:(EU-Grant)945539},
typ = {PUB:(DE-HGF)3 / PUB:(DE-HGF)11},
urn = {urn:nbn:de:0001-20231004081300012-6971752-0},
doi = {10.34734/FZJ-2023-03187},
url = {https://juser.fz-juelich.de/record/1010679},
}