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@ARTICLE{ffing:1026361,
author = {Üffing, Alina and Gold, Lisa and Gensch, Thomas and
Weiergräber, Oliver H. and Hoffmann, Silke and Willbold,
Dieter},
title = {{H}ighlighting the hidden: monitoring the avidity-driven
association of a fluorescent {GABARAP} tandem with
microtubules in living cells},
journal = {Autophagy reports},
volume = {3},
number = {1},
issn = {2769-4127},
address = {London},
publisher = {Taylor $\&$ Francis Group},
reportid = {FZJ-2024-03392},
pages = {2348899},
year = {2024},
note = {This is an Open Access article distributed under the terms
of the Creative Commons Attribution License
(http://creativecommons.org/licenses/by/4.0/), which permits
unrestricted use, distribution, and reproduction in any
medium, provided the original work is properly cited. The
terms on which this article has been published allow the
posting of the Accepted Manuscript in a repository by the
author(s) or with their consent.},
abstract = {GABARAP, like other ATG8 proteins, is a ubiquitin-like
modifier and its C-terminal lipid conjugation enables
association with cellular membranes. To prevent interference
with the lipidation process, N-terminal fluorescent protein
(FP) tagging strategies have become the standard for
studying ATG8 localization and function in living cells,
significantly contributing to our understanding of this
protein family’s multifaceted roles. We employed live cell
imaging with particular emphasis on a GABARAP split-tandem
construct, GABARAP(G116A)-mTagBFP2-GABARAP (G-B-G), which
retains both a free N-terminus and a lipidation-competent
c-terminus, while bivalence creates a gain in affinity
conferred by avidity. Notably, reminiscent of early in vitro
studies demonstrating an interaction of GABARAP and tubulin,
our results revealed a robust association of G-B-G with the
microtubule network in living cells. We show that the
presence of several basic residues in the amino-terminal
helical subdomain of GABARAP and avidity emerged as
essential for robust MT association, whereas lipidation
ability was not decisive. Interestingly, while the position
of the FP-tag had little influence on the result, the nature
of the FP itself was crucial, with mTagBFP2 being required
for tracking GABARAP tandems in the vicinity of MTs. Though
artificial effects cannot be excluded, we assume that G-B-G,
with its increased avidity, can give visibility to processes
that are based on inherently weak interactions, and thus can
help elucidate potential roles of GABARAP e.g. in
microtubule-associated processes that are integral to
autophagy-related and -unrelated cellular transport.},
cin = {IBI-7 / IBI-1},
ddc = {610},
cid = {I:(DE-Juel1)IBI-7-20200312 / I:(DE-Juel1)IBI-1-20200312},
pnm = {5241 - Molecular Information Processing in Cellular Systems
(POF4-524) / SFB 1208 B02 - Spezifische Rollen von Atg8s im
Vesikeltransport (B02) (289554527)},
pid = {G:(DE-HGF)POF4-5241 / G:(GEPRIS)289554527},
typ = {PUB:(DE-HGF)16},
doi = {10.1080/27694127.2024.2348899},
url = {https://juser.fz-juelich.de/record/1026361},
}