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@ARTICLE{Sarid:139281,
author = {Sarid, L. and Feldmeyer, D. and Gidon, A. and Sakmann, B.
and Segev, I.},
title = {{C}ontribution of {I}ntracolumnar {L}ayer 2/3-to-{L}ayer
2/3 {E}xcitatory {C}onnections in {S}haping the {R}esponse
to {W}hisker {D}eflection in {R}at {B}arrel {C}ortex},
journal = {Cerebral cortex},
volume = {25},
number = {4},
issn = {1460-2199},
address = {Oxford},
publisher = {Oxford Univ. Press},
reportid = {FZJ-2013-05280},
pages = {849-858},
year = {2015},
abstract = {This computational study integrates anatomical and
physiological data to assess the functional role of the
lateral excitatory connections between layer 2/3 (L2/3)
pyramidal cells (PCs) in shaping their response during early
stages of intracortical processing of a whisker deflection
(WD). Based on in vivo and in vitro recordings, and 3D
reconstructions of connected pairs of L2/3 PCs, our model
predicts that: 1) AMPAR and NMDAR conductances/synapse are
0.52 ± 0.24 and 0.40 ± 0.34 nS, respectively; 2) following
WD, connection between L2/3 PCs induces a composite EPSPs of
7.6 ± 1.7 mV, well below the threshold for action potential
(AP) initiation; 3) together with the excitatory feedforward
L4-to-L2/3 connection, WD evoked a composite EPSP of 16.3 ±
3.5 mV and a probability of 0.01 to generate an AP. When
considering the variability in L4 spiny neurons
responsiveness, it increased to 17.8 ± 11.2 mV; this 3-fold
increase in the SD yielded AP probability of 0.35; 4) the
interaction between L4-to-L2/3 and L2/3-to-L2/3 inputs is
highly nonlinear; 5) L2/3 dendritic morphology significantly
affects L2/3 PCs responsiveness. We conclude that early
stages of intracortical signaling of WD are dominated by a
combination of feedforward L4–L2/3 and L2/3–L2/3 lateral
connections.},
cin = {INM-2 / JARA-BRAIN},
ddc = {610},
cid = {I:(DE-Juel1)INM-2-20090406 / $I:(DE-82)080010_20140620$},
pnm = {571 - Connectivity and Activity (POF3-571)},
pid = {G:(DE-HGF)POF3-571},
typ = {PUB:(DE-HGF)16},
UT = {WOS:000353838600002},
doi = {10.1093/cercor/bht268},
url = {https://juser.fz-juelich.de/record/139281},
}