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@ARTICLE{Inoue:16351,
author = {Inoue, R. and Biehl, R. and Rosenkranz, T. and Fitter, J.
and Monkenbusch, M. and Radulescu, A. and Farago, B. and
Richter, D.},
title = {{L}arge domain fluctuations on 50ns timescale enable
catalytic activity in phoshpglycerate kinase},
journal = {Biophysical journal},
volume = {99},
number = {7},
issn = {0006-3495},
address = {New York, NY},
publisher = {Rockefeller Univ. Press},
reportid = {PreJuSER-16351},
pages = {2309 - 2317},
year = {2010},
note = {T.R. acknowledges financial support by the International
Helmholtz Research School on Biophysics and Soft Matter
(BioSoft).},
abstract = {Large-scale domain motions of enzymes are often essential
for their biological function. Phosphoglycerate kinase has a
wide open domain structure with a hinge near the active
center between the two domains. Applying neutron spin echo
spectroscopy and small-angle neutron scattering we have
investigated the internal domain dynamics. Structural
analysis reveals that the holoprotein in solution seems to
be more compact compared to the crystal structure but would
not allow the functionally important phosphoryl transfer
between the substrates if the protein were static. Brownian
large-scale domain fluctuation dynamics on a timescale of 50
ns was revealed by neutron spin echo spectroscopy. The
dynamics observed was compared to the displacement patterns
of low-frequency normal modes. The displacements along the
normal-mode coordinates describe our experimental results
reasonably well. In particular, the domain movements
facilitate a close encounter of the key residues in the
active center to build the active configuration. The
observed dynamics shows that the protein has the flexibility
to allow fluctuations and displacements that seem to enable
the function of the protein. Moreover, the presence of the
substrates increases the rigidity, which is deduced from a
faster dynamics with smaller amplitude.},
keywords = {Biocatalysis / Diffusion / Kinetics / Models, Molecular /
Neutron Diffraction / Phosphoglycerate Kinase: chemistry /
Phosphoglycerate Kinase: metabolism / Protein Structure,
Secondary / Protein Structure, Tertiary / Saccharomyces
cerevisiae: enzymology / Scattering, Small Angle /
Structure-Activity Relationship / Time Factors /
Phosphoglycerate Kinase (NLM Chemicals) / J (WoSType)},
cin = {ICS-1 / JCNS-1 / JCNS (München) ; Jülich Centre for
Neutron Science JCNS (München) ; JCNS-FRM-II / ICS-5},
ddc = {570},
cid = {I:(DE-Juel1)ICS-1-20110106 / I:(DE-Juel1)JCNS-1-20110106 /
I:(DE-Juel1)JCNS-FRM-II-20110218 /
I:(DE-Juel1)ICS-5-20110106},
pnm = {BioSoft: Makromolekulare Systeme und biologische
Informationsverarbeitung / Großgeräte für die Forschung
mit Photonen, Neutronen und Ionen (PNI)},
pid = {G:(DE-Juel1)FUEK505 / G:(DE-Juel1)FUEK415},
experiment = {EXP:(DE-MLZ)KWS1-20140101},
shelfmark = {Biophysics},
typ = {PUB:(DE-HGF)16},
pubmed = {pmid:20923666},
pmc = {pmc:PMC3042550},
UT = {WOS:000282850600037},
doi = {10.1016/j.bpj.2010.08.017},
url = {https://juser.fz-juelich.de/record/16351},
}