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@ARTICLE{Zerrad:16685,
author = {Zerrad, L. and Merli, A. and Schröder, G.F. and Varga, A.
and Gráczer, E. and Pernot, P. and Round, A. and Vas, M.
and Bowler, M.W.},
title = {{A} {S}pring {L}oaded {R}elease {M}echanism {R}egulates
{D}omain {M}ovement and {C}atalysis in {P}hosphoglycerate
{K}inase.},
journal = {The journal of biological chemistry},
volume = {286},
issn = {0021-9258},
address = {Bethesda, Md.},
publisher = {Soc.},
reportid = {PreJuSER-16685},
pages = {14040 - 14048},
year = {2011},
note = {This work was supported in part by Hungarian National
Research Grant 77978.},
abstract = {Phosphoglycerate kinase (PGK) is the enzyme responsible for
the first ATP-generating step of glycolysis and has been
implicated extensively in oncogenesis and its development.
Solution small angle x-ray scattering (SAXS) data, in
combination with crystal structures of the enzyme in complex
with substrate and product analogues, reveal a new
conformation for the resting state of the enzyme and
demonstrate the role of substrate binding in the preparation
of the enzyme for domain closure. Comparison of the x-ray
scattering curves of the enzyme in different states with
crystal structures has allowed the complete reaction cycle
to be resolved both structurally and temporally. The enzyme
appears to spend most of its time in a fully open
conformation with short periods of closure and catalysis,
thereby allowing the rapid diffusion of substrates and
products in and out of the binding sites. Analysis of the
open apoenzyme structure, defined through deformable elastic
network refinement against the SAXS data, suggests that
interactions in a mostly buried hydrophobic region may favor
the open conformation. This patch is exposed on domain
closure, making the open conformation more thermodynamically
stable. Ionic interactions act to maintain the closed
conformation to allow catalysis. The short time PGK spends
in the closed conformation and its strong tendency to rest
in an open conformation imply a spring-loaded release
mechanism to regulate domain movement, catalysis, and
efficient product release.},
keywords = {Adenosine Triphosphate: chemistry / Amino Acid Sequence /
Animals / Binding Sites / Biophysics: methods / Catalysis /
Crystallography, X-Ray: methods / Humans / Mice / Molecular
Sequence Data / Phosphoglycerate Kinase: chemistry / Protein
Conformation / Protein Structure, Secondary / Protein
Structure, Tertiary / Scattering, Radiation / Sequence
Homology, Amino Acid / Thermodynamics / Adenosine
Triphosphate (NLM Chemicals) / Phosphoglycerate Kinase (NLM
Chemicals) / J (WoSType)},
cin = {ICS-6},
ddc = {570},
cid = {I:(DE-Juel1)ICS-6-20110106},
pnm = {Funktion und Dysfunktion des Nervensystems / BioSoft:
Makromolekulare Systeme und biologische
Informationsverarbeitung},
pid = {G:(DE-Juel1)FUEK409 / G:(DE-Juel1)FUEK505},
shelfmark = {Biochemistry $\&$ Molecular Biology},
typ = {PUB:(DE-HGF)16},
pubmed = {pmid:21349853},
pmc = {pmc:PMC3077604},
UT = {WOS:000289556200026},
doi = {10.1074/jbc.M110.206813},
url = {https://juser.fz-juelich.de/record/16685},
}