TY - JOUR AU - Galkin, V.E. AU - Orlova, A. AU - Kudryashov, D.S. AU - Solodukhin, A. AU - Reisler, E. AU - Schröder, G.F. AU - Egelman, E.H. TI - Remodeling of actin filaments by ADF/cofilin proteins JO - Proceedings of the National Academy of Sciences of the United States of America VL - 108 SN - 0027-8424 CY - Washington, DC PB - Academy M1 - PreJuSER-19044 SP - 20568 - 20572 PY - 2011 N1 - This work was supported by National Institutes of Health Grants GM081303 (E. H. E.) and GM077190 (E.R.). AB - Cofilin/ADF proteins play key roles in the dynamics of actin, one of the most abundant and highly conserved eukaryotic proteins. We used cryoelectron microscopy to generate a 9-Å resolution three-dimensional reconstruction of cofilin-decorated actin filaments, the highest resolution achieved for a complex of F-actin with an actin-binding protein. We show that the cofilin-induced change in the filament twist is due to a unique conformation of the actin molecule unrelated to any previously observed state. The changes between the actin protomer in naked F-actin and in the actin-cofilin filament are greater than the conformational changes between G- and F-actin. Our results show the structural plasticity of actin, suggest that other actin-binding proteins may also induce large but different conformational changes, and show that F-actin cannot be described by a single molecular model. KW - Actin Depolymerizing Factors: chemistry KW - Actins: chemistry KW - Cofilin 2: chemistry KW - Cryoelectron Microscopy: methods KW - Cytoskeleton: chemistry KW - Gene Library KW - Humans KW - Microscopy, Electron: methods KW - Models, Molecular KW - Molecular Conformation KW - Muscle, Skeletal: metabolism KW - Polymers: chemistry KW - Protein Conformation KW - Protein Structure, Secondary KW - Actin Depolymerizing Factors (NLM Chemicals) KW - Actins (NLM Chemicals) KW - Cofilin 2 (NLM Chemicals) KW - Polymers (NLM Chemicals) KW - J (WoSType) LB - PUB:(DE-HGF)16 C6 - pmid:22158895 C2 - pmc:PMC3251117 UR - <Go to ISI:>//WOS:000298289400064 DO - DOI:10.1073/pnas.1110109108 UR - https://juser.fz-juelich.de/record/19044 ER -