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@ARTICLE{Voiniciuc:203204,
author = {Voiniciuc, Catalin and Schmidt, Maximilian Heinrich-Wilhelm
and Berger, Adeline and Yang, Bo and Ebert, Berit and
Scheller, Henrik Vibe and North, Helen M. and Usadel, Björn
and Günl, Markus},
title = {{MUCI}10 {P}roduces {G}alactoglucomannan {T}hat {M}aintains
{P}ectin and {C}ellulose {A}rchitecture in {A}rabidopsis
{S}eed {M}ucilage},
journal = {Plant physiology},
volume = {169},
number = {1},
issn = {1532-2548},
address = {Rockville, Md.},
publisher = {Soc.},
reportid = {FZJ-2015-05202},
pages = {pp.00851.2015 -},
year = {2015},
abstract = {Plants invest a lot of their resources into the production
of an extracellular matrix built of polysaccharides. While
the composition of the cell wall is relatively well
characterized, the functions of the individual polymers and
the enzymes that catalyze their biosynthesis remain poorly
understood. We exploited the Arabidopsis thaliana seed coat
epidermis (SCE) to study cell wall synthesis. SCE cells
produce mucilage, a specialized secondary wall that is rich
in pectin, at a precise stage of development. A
co-expression search for MUCILAGE-RELATED (MUCI) genes
identified MUCI10 as a key determinant of mucilage
properties. MUCI10, a member of the GT34 family, is closely
related to a fenugreek enzyme that has in vitro
galactomannan α-1,6-galactosyltransferase activity. Our
detailed analysis of the muci10 mutants demonstrates that
mucilage contains highly branched galactoglucomannan (GGM)
rather than unbranched glucomannan. MUCI10 likely decorates
glucomannan, synthesized by CSLA2, with galactose residues
in vivo. The degree of galactosylation is essential for the
synthesis of the GGM backbone, the structure of cellulose,
mucilage density, as well as the adherence of pectin. We
propose that GGM scaffolds control mucilage architecture
along with cellulosic rays, and show that Arabidopsis SCE
cells represent an excellent model to study the synthesis
and function of GGM. Arabidopsis natural varieties with
defects similar to muci10 mutants may reveal additional
genes involved in GGM synthesis. Since GGM is the most
abundant hemicellulose in the secondary walls of
gymnosperms, understanding its biosynthesis may facilitate
improvements in the production of valuable commodities from
softwoods.},
cin = {IBG-2},
ddc = {580},
cid = {I:(DE-Juel1)IBG-2-20101118},
pnm = {582 - Plant Science (POF3-582)},
pid = {G:(DE-HGF)POF3-582},
typ = {PUB:(DE-HGF)16},
UT = {WOS:000360930600033},
pubmed = {pmid:26220953},
doi = {10.1104/pp.15.00851},
url = {https://juser.fz-juelich.de/record/203204},
}