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@ARTICLE{Stadler:23114,
author = {Stadler, A.M. and Garvey, C.J. and Bocahut, A. and
Sacquin-Mora, S. and Digel, I. and Schneider, G.J. and
Natali, F. and Artmann, G.M. and Zaccai, G.},
title = {{T}hermal fluctuations of haemoglobin from different
species: adaptation to temperature via conformational
dynamics},
journal = {Interface},
volume = {9},
issn = {1742-5689},
address = {London},
publisher = {The Royal Society},
reportid = {PreJuSER-23114},
pages = {2845 - 2855},
year = {2012},
note = {Platypus RBC was provided by Nick Gust (Department of
Primary Industries Tasmania, Australia) and Dominic Geraghty
(University of Tasmania, Australia). Crocodile blood was
provided by Crocodylus Park (Darwin, Australia). We thank
Prof. Philip Kuchel (University of Sydney, Australia) for
the use of his laboratory to purify crocodile and platypus
Hb. Chicken blood was provided by Mario Suarez Avello and
Josefa Garcia Alvarez. I. D. was sponsored by the BMBF
Programme 'FHProfUnd' grant no. 1736X08. We also thank the
Ministry of Innovation Science and Research of the federal
state North Rhine-Westphalia for supporting Prof. Aysegul
(Temiz) Artmann's project Dynamischer Strukturubergang bei
Korpertemperaturen (FKZ: 800 009 04). A. M. S. thanks Prof.
Georg Buldt and Prof. Dieter Richter for continuous support.
This work is based on experiments performed at the Institute
Laue-Langevin (ILL), Grenoble, France, and at the
Forschungs-Neutronenquelle Heinz Maier-Leibnitz (FRM II),
Garching, Germany.},
abstract = {Thermodynamic stability, configurational motions and
internal forces of haemoglobin (Hb) of three endotherms
(platypus, Ornithorhynchus anatinus; domestic chicken,
Gallus gallus domesticus and human, Homo sapiens) and an
ectotherm (salt water crocodile, Crocodylus porosus) were
investigated using circular dichroism, incoherent elastic
neutron scattering and coarse-grained Brownian dynamics
simulations. The experimental results from Hb solutions
revealed a direct correlation between protein resilience,
melting temperature and average body temperature of the
different species on the 0.1 ns time scale. Molecular forces
appeared to be adapted to permit conformational fluctuations
with a root mean square displacement close to 1.2 Å at the
corresponding average body temperature of the endotherms.
Strong forces within crocodile Hb maintain the amplitudes of
motion within a narrow limit over the entire temperature
range in which the animal lives. In fully hydrated powder
samples of human and chicken, Hb mean square displacements
and effective force constants on the 1 ns time scale showed
no differences over the whole temperature range from 10 to
300 K, in contrast to the solution case. A complementary
result of the study, therefore, is that one hydration layer
is not sufficient to activate all conformational
fluctuations of Hb in the pico- to nanosecond time scale
which might be relevant for biological function.
Coarse-grained Brownian dynamics simulations permitted to
explore residue-specific effects. They indicated that
temperature sensing of human and chicken Hb occurs mainly at
residues lining internal cavities in the β-subunits.},
keywords = {J (WoSType)},
cin = {ICS-1 / JCNS-1 / JCNS (München) ; Jülich Centre for
Neutron Science JCNS (München) ; JCNS-FRM-II},
ddc = {500},
cid = {I:(DE-Juel1)ICS-1-20110106 / I:(DE-Juel1)JCNS-1-20110106 /
I:(DE-Juel1)JCNS-FRM-II-20110218},
pnm = {BioSoft: Makromolekulare Systeme und biologische
Informationsverarbeitung / 544 - In-house Research with PNI
(POF2-544)},
pid = {G:(DE-Juel1)FUEK505 / G:(DE-HGF)POF2-544},
experiment = {EXP:(DE-MLZ)SPHERES-20140101},
shelfmark = {Multidisciplinary Sciences},
typ = {PUB:(DE-HGF)16},
pubmed = {pmid:22696485},
pmc = {pmc:PMC3479923},
UT = {WOS:000309269100010},
doi = {10.1098/rsif.2012.0364},
url = {https://juser.fz-juelich.de/record/23114},
}