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@PHDTHESIS{Unthan:256301,
author = {Unthan, Simon},
title = {{R}obot-{A}ssisted {P}henotyping of {G}enome-{R}educed
${C}orynebacterium$ $glutamicum$ {S}train {L}ibraries to
{D}raft a {C}hassis {O}rganism},
volume = {132},
school = {RWTH Aachen},
type = {Dr.},
address = {Jülich},
publisher = {Forschungszentrum Jülich GmbH Zentralbibliothek, Verlag},
reportid = {FZJ-2015-06265},
isbn = {978-3-95806-169-9},
series = {Schriften des Forschungszentrums Jülich. Reihe
Schlüsseltechnologien / Key Technologies},
pages = {122 S.},
year = {2016},
note = {RWTH Aachen, Diss., 2015},
abstract = {In this work, concepts were developed and applied to guide
the construction of a $\textit{Corynebacterium glutamicum}$
chassis organism for synthetic biology approaches. The aim
was to delete irrelevant genes from the wild type strain in
order to obtain a chassis growing on defined CGXII medium
with D-glucose with unaltered biological fitness, which was
defined by the maximum specific growth rate and biomass
yield. Initially, workflows were developed on a robotic Mini
Pilot Plant (MPP), for example, to harvest cell-free
cultivation supernatants from BioLector cultivations in
response to individually defined triggers. Subsequently,
assays for amino acids and D-glucose were established in
384-well plate scale in order to quantify these metabolites
in cell-free culture supernatants in fully automated
workflows [1]. During initial reference experiments,
protocatechuic acid was identified as a hidden co-substrate
in the well-known defined CGXII medium. The additional TCA
feed via acetyl-CoA and succinyl-CoA, which are derived from
protocatechuic acid, elevates the growth rate by about 50 \%
in highly diluted cultures [2]. The first step toward a
chassis was the deletion of prophage elements contributing
to about 6.7 \% of the $\textit{C. glutamicum}$ genome. The
respective strain MB001 showed unaltered biological fitness
and an increased heterologous protein expression, caused by
the removal of a restriction-modification system in prophage
CGP3 [3]. As a next step, 36 strains with deletion of
non-essential gene clusters were tested thoroughly and 26
clusters were found irrelevant for the biological fitness of
$\textit{C. glutamicum}$ and offered the potential to reduce
the genome by about 22 \% [4]. Some clusters were also
deleted in the L-lysine model producer DM1933 and the
derived strain GRLP45 showed an 51 \% increased L-lysine
titer applying the automated MPP methods, what was finally
confirmed in lab-scale bioreactors [1]. During the final
combinatorial deletion of irrelevant gene clusters, some
interdependencies were observed resulting in a decreased of
biological fitness of the respective strains. One of those
strains was characterized in-depth and revealed the general
interplay of ribosome capacity and maximum growth rate of
$\textit{C. glutamicum}$. In the end, two pre-chassis,
namely W127 and W121, were obtained that displayed a total
genome reduction of 8.8 \% and 12.8 \%, respectively. Both
strains fulfilled the target criteria of unaltered
biological fitness on defined CGXII medium in BioLector
cultivations. Finally, the in-depth analysis of both
pre-chassis in bioreactors revealed a morphological
divergence of W121 which could be narrowed down to a single
cluster deletion. However, W127 did not show any drawback
compared to the wild type when tested under stress
conditions and on different cultivation scales. In fact,
this strain even grew faster on some C-sources, making it a
good basis for synthetic biology approaches.},
cin = {IBG-1},
cid = {I:(DE-Juel1)IBG-1-20101118},
pnm = {89581 - Biotechnology (POF2-89581)},
pid = {G:(DE-HGF)POF2-89581},
typ = {PUB:(DE-HGF)3 / PUB:(DE-HGF)11},
urn = {urn:nbn:de:0001-2017040716},
url = {https://juser.fz-juelich.de/record/256301},
}