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@ARTICLE{Stadler:283547,
author = {Stadler, Andreas and Knieps-Grünhagen, Esther and Bocola,
Marco and Lohstroh, Wiebke and Zamponi, Michaela and Krauss,
Ulrich},
title = {{P}hotoactivation {R}educes {S}ide-{C}hain {D}ynamics of a
{LOV} {P}hotoreceptor},
journal = {Biophysical journal},
volume = {110},
number = {5},
issn = {0006-3495},
address = {Cambridge, Mass.},
publisher = {Cell Press},
reportid = {FZJ-2016-01863},
pages = {1064 - 1074},
year = {2016},
abstract = {We used neutron-scattering experiments to probe the
conformational dynamics of the light, oxygen, voltage (LOV)
photoreceptor PpSB1-LOV from Pseudomonas putida in both the
dark and light states. Global protein diffusion and internal
macromolecular dynamics were measured using incoherent
neutron time-of-flight and backscattering spectroscopy on
the picosecond to nanosecond timescales. Global protein
diffusion of PpSB1-LOV is not influenced by photoactivation.
Observation-time-dependent global diffusion coefficients
were found, which converge on the nanosecond timescale
toward diffusion coefficients determined by dynamic light
scattering. Mean-square displacements of localized internal
motions and effective force constants, <k′>, describing
the resilience of the proteins were determined on the
respective timescales. Photoactivation significantly
modifies the flexibility and the resilience of PpSB1-LOV. On
the fast, picosecond timescale, small changes in the
mean-square displacement and <k′> are observed, which are
enhanced on the slower, nanosecond timescale.
Photoactivation results in a slightly larger resilience of
the photoreceptor on the fast, picosecond timescale, whereas
in the nanosecond range, a significantly less resilient
structure of the light-state protein is observed. For a
residue-resolved interpretation of the experimental
neutron-scattering data, we analyzed molecular dynamics
simulations of the PpSB1-LOV X-ray structure. Based on these
data, it is tempting to speculate that light-induced changes
in the protein result in altered side-chain mobility mostly
for residues on the protruding Jα helix and on the LOV-LOV
dimer interface. Our results provide strong experimental
evidence that side-chain dynamics play a crucial role in
photoactivation and signaling of PpSB1-LOV via modulation of
conformational entropy.},
cin = {ICS-1 / Neutronenstreuung ; JCNS-1 / JCNS (München) ;
Jülich Centre for Neutron Science JCNS (München) ;
JCNS-FRM-II / IMET},
ddc = {570},
cid = {I:(DE-Juel1)ICS-1-20110106 / I:(DE-Juel1)JCNS-1-20110106 /
I:(DE-Juel1)JCNS-FRM-II-20110218 /
I:(DE-Juel1)IMET-20090612},
pnm = {551 - Functional Macromolecules and Complexes (POF3-551) /
6G4 - Jülich Centre for Neutron Research (JCNS) (POF3-623)
/ 6215 - Soft Matter, Health and Life Sciences (POF3-621)},
pid = {G:(DE-HGF)POF3-551 / G:(DE-HGF)POF3-6G4 /
G:(DE-HGF)POF3-6215},
experiment = {EXP:(DE-MLZ)SPHERES-20140101 /
EXP:(DE-MLZ)TOF-TOF-20140101},
typ = {PUB:(DE-HGF)16},
UT = {WOS:000371850000015},
pubmed = {pmid:26958884},
doi = {10.1016/j.bpj.2016.01.021},
url = {https://juser.fz-juelich.de/record/283547},
}