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@PHDTHESIS{Pieruschka:45486,
author = {Pieruschka, Roland},
title = {{E}ffect of internal leaf structures on gas exchange of
leaves},
volume = {56},
school = {Universität Düsseldorf},
type = {Dr. (Univ.)},
address = {Jülich},
publisher = {Forschungszentrum Jülich GmbH Zentralbibliothek, Verlag},
reportid = {PreJuSER-45486},
isbn = {3-89336-403-X},
series = {Schriften des Forschungszentrums Jülich. Reihe Umwelt /
Environment},
pages = {120 S.},
year = {2005},
note = {Record converted from VDB: 12.11.2012; Universität
Düsseldorf, Diss., 2005},
abstract = {Gas exchange of leaves is generally considered as the
interchange of gaseous compounds between the leaf interior
and ambient air. Once inside the leaf, CO$_{2}$ can diffuse
along its concentration gradients mainly regarded in the
vertical direction of the blade towards the assimilating
tissues. Lateral gas diffusion within intercellular air
spaces may be much more effective than has been considered
so far which depends on anatomical features of leaves. In
heterobaric leaves, lateral diffusion is restricted by
bundle-sheath extensions and the mesophyll is composed of
closed compartments. Homobaric leaves, however, lack such
extensions and the leaves have large interconnected
intercellular air spaces. The specific internal gas
diffusion properties of the leaves were characterized by gas
conductivities. Gas conductivity was larger in lateral than
in the vertical direction of homobaric leaf blades. However,
there was a large variability of the size and property of
the intercellular air space among different species. When
`clamp-on´ leaf chambers were used it was found that
lateral diffusion inside leaves seriously affected gas
exchange measurements. The impact of lateral CO$_{2}$
diffusion on gas exchange measurement was substantial when
exchange rates were low. Homobaric leaves showed internal
lateral gas fluxes when an overpressure was applied to the
leaf chamber which has been used in commercial gas exchange
systems to minimise the effects of leaks in the leaf
chamber. It was found here that overpressure affected
CO$_{2}$ and H$_{2}$O exchange rates of homobaric leaves
substantially larger than the theoretical direct impact of
air pressure on gas exchange processes. Gas gradients inside
leaves emerged when a leaf part was shaded and the adjacent
area of the leaf blade illuminated. Respiratory CO$_{2}$
evolved in the shaded region diffused to the illuminated
area were it was fixed by photosynthesis. These processes
obviously increased the photosynthetic efficiency along the
light/shade borderline as was visualized by chlorophyll
fluorescence imaging techniques. The recycling of
respiratory CO$_{2}$ from distant shaded areas was found to
be larger when stomatal conductance was low as is the case
under drought stress. Thus, when a homobaric leaf was
illuminated by lightflecks, additional CO$_{2}$ increased
the carbon gain, water use efficiency, and reduced light
stress. It was hypothesized that homobaric leaf anatomy is a
trait which has evolved under certain environmental
conditions.},
cin = {ICG-III},
ddc = {333.7},
cid = {I:(DE-Juel1)VDB49},
pnm = {Chemie und Dynamik der Geo-Biosphäre},
pid = {G:(DE-Juel1)FUEK257},
typ = {PUB:(DE-HGF)11 / PUB:(DE-HGF)3},
url = {https://juser.fz-juelich.de/record/45486},
}