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@ARTICLE{Janzik:45605,
author = {Janzik, I. and Preiskowski, S. and Kneifel, H.},
title = {{O}zone has dramatic effects on the regulation of the
prechorismate pathway in tobacco ({N}icotiana tabacum
{L}.cv. {B}el {W}3)},
journal = {Planta},
volume = {223},
issn = {0032-0935},
address = {Berlin},
publisher = {Springer},
reportid = {PreJuSER-45605},
pages = {20 - 27},
year = {2005},
note = {Record converted from VDB: 12.11.2012},
abstract = {The accumulation of aromatic secondary metabolites is a
well-known element of the plant response to ozone. Most of
these metabolites are synthesized via the three aromatic
amino acids phenylalanine, tyrosine and tryptophan. Before
branching, the biosynthetic pathway to the three amino acids
shares seven enzymatic steps, called the prechorismate
pathway, catalysed by 3-deoxy-D:
-arabino-heptulosonate-7-phosphate (DAHP) synthase [EC
2.5.1.54], 3-dehydroquinate synthase [EC 4.2.3.4],
3-dehydroquinate dehydratase [EC 4.2.1.10]-shikimate
5-dehydrogenase [EC 1.1.1.25], shikimate kinase [EC
2.7.1.71], 5-enolpyruvylshikimate 3-phosphate synthase [EC
2.5.1.19] and chorismate synthase [EC 4.2.3.5]). We have
studied the transcript level of these enzymes and the
aromatic metabolite profile in the ozone sensitive tobacco
cultivar BelW3 (Nicotiana tabacum L. cv Bel W3), when
exposed to an acute ozone pulse (160 nl l(-1), 5 h).
Specific cDNA-fragments of the corresponding six genes were
isolated from tobacco Bel W3 and used as probes for
determining the expression of the prechorismate pathway
genes. The fully expanded leaves of ozone treated plants,
which developed symptoms like necrotic leaf spots and
accumulation of aromatic metabolites, showed a clear
induction of the shikimate pathway genes; indicating, that
this induction is linked to the development of the symptoms.
Distinct kinetics and magnitudes were observed in tobacco
leaves for the ozone dependent enhanced mRNA accumulation of
the aforementioned genes in BelW3. The strongest and
earliest induction due to ozone treatment could be observed
for DAHP synthase. An isoform-specific analysis of the
transcripts showed a strong induction on transcript level
only for one of three isoforms, which was followed by the
induction of the DAHP synthase also on protein level. The
different induction kinetics of the prechorismate pathway
genes indicate that their regulation in response to ozone
might be regulated by different signals, for example,
ethylene, reactive oxygen species or salicylic acid, which
also occur with different kinetics and thus may play
different roles in the plant response to ozone.},
keywords = {3-Deoxy-7-Phosphoheptulonate Synthase / Aldehyde-Lyases:
biosynthesis / Aldehyde-Lyases: genetics / Amino Acids,
Aromatic: biosynthesis / Blotting, Northern / Chorismic
Acid: metabolism / Chromatography, High Pressure Liquid /
DNA, Complementary / DNA, Plant / Enzyme Induction / Gene
Expression Regulation, Enzymologic / Gene Expression
Regulation, Plant / Ozone: pharmacology /
Phosphotransferases (Alcohol Group Acceptor): metabolism /
Plant Leaves / Tobacco: enzymology / Tobacco: genetics /
Tobacco: metabolism / Amino Acids, Aromatic (NLM Chemicals)
/ DNA, Complementary (NLM Chemicals) / DNA, Plant (NLM
Chemicals) / Ozone (NLM Chemicals) / Chorismic Acid (NLM
Chemicals) / 3-Deoxy-7-Phosphoheptulonate Synthase (NLM
Chemicals) / Phosphotransferases (Alcohol Group Acceptor)
(NLM Chemicals) / shikimate kinase (NLM Chemicals) /
Aldehyde-Lyases (NLM Chemicals) / J (WoSType)},
cin = {ICG-III},
ddc = {580},
cid = {I:(DE-Juel1)VDB49},
pnm = {Chemie und Dynamik der Geo-Biosphäre},
pid = {G:(DE-Juel1)FUEK257},
shelfmark = {Plant Sciences},
typ = {PUB:(DE-HGF)16},
pubmed = {pmid:16078071},
UT = {WOS:000233725000003},
doi = {10.1007/s00425-005-0060-8},
url = {https://juser.fz-juelich.de/record/45605},
}