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@ARTICLE{Akola:51046,
author = {Akola, J. and Jones, R. O.},
title = {{D}ensity functional calculations of {ATP} systems {II}:
{ATP} hydrolysis at the active site of actin},
journal = {The journal of physical chemistry / B},
volume = {110},
issn = {1520-6106},
address = {Washington, DC},
publisher = {Soc.},
reportid = {PreJuSER-51046},
pages = {8121 - 8129},
year = {2006},
note = {Record converted from VDB: 12.11.2012},
abstract = {The hydrolysis of adenosine 5'-triphosphate (ATP) at the
active site of actin has been studied using density
functional calculations. The active site is modeled by the
triphosphate tail of ATP, an Mg cation, surrounding water
molecules, and the nearby protein residues. Four reaction
paths have been followed by constraining coordinates that
allow phosphate stretching, nucleophilic attack of the
catalytic water, and OH(-) formation via water
deprotonation. The lowest-energy barrier (21.0 kcal/mol) is
obtained for a dissociative reaction where the terminal
phosphate breaks on approaching the catalytic water,
followed by proton release via a proton wire mechanism. A
higher barrier (39.6 kcal/mol) results for an associative
reaction path where OH(-) is formed first, with a
pentacoordinated phosphorus atom (P-O distances 2.1 A).
Stretching the terminal bridging P-O bond results in bond
rupture at 2.8 A with an energy barrier of 28.8 kcal/mol.
The residues Gln137 and His161 are not important in the
reactions, but insight into their roles in vivo has been
obtained. The favored coordination of the end products
H(2)PO(4)(-) and ADP(3-) includes a hydrogen bond and an
O-Mg-O bridge between the phosphates as well as a hydrogen
bond between H(2)PO(4)(-) and the Ser14 side chain. The
total energy is 2.1 kcal/mol lower than in the initial
reactants. Classical simulations of ATP- and ADP.P(i)-actin
show few hydrolysis-induced differences in the protein
structure, indicating that phosphate migration is necessary
for a change in conformation.},
keywords = {Actins: chemistry / Adenosine Triphosphate: chemistry /
Binding Sites / Hydrogen Bonding / Hydrolysis / Models,
Molecular / Saccharomyces cerevisiae: chemistry / Actins
(NLM Chemicals) / Adenosine Triphosphate (NLM Chemicals) / J
(WoSType)},
cin = {IFF-TH-I},
ddc = {530},
cid = {I:(DE-Juel1)VDB30},
pnm = {Kondensierte Materie},
pid = {G:(DE-Juel1)FUEK414},
shelfmark = {Chemistry, Physical},
typ = {PUB:(DE-HGF)16},
pubmed = {pmid:16610915},
UT = {WOS:000236992100069},
doi = {10.1021/jp054921d},
url = {https://juser.fz-juelich.de/record/51046},
}