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@ARTICLE{Kottke:51338,
author = {Kottke, T. and Ahmad, M. and Batschauer, A. and Heberle,
J.},
title = {{B}lue-{L}ight-{I}nduced {C}hanges in {A}rabidopsis
{C}ryptochrome 1 {P}robed by {FTIR} {D}ifference
{S}pectroscopy},
journal = {Biochemistry},
volume = {45},
issn = {0006-2960},
address = {Columbus, Ohio},
publisher = {American Chemical Society},
reportid = {PreJuSER-51338},
pages = {2472 - 2479},
year = {2006},
note = {Record converted from VDB: 12.11.2012},
abstract = {Cryptochromes are blue-light photoreceptors that regulate a
variety of responses in animals and plants, including
circadian entrainment in Drosophila and photomorphogenesis
in Arabidopsis. They comprise a photolyase homology region
(PHR) of about 500 amino acids and a C-terminal extension of
varying length. In the PHR domain, flavin adenine
dinucleotide (FAD) is noncovalently bound. The presence of a
second chromophore, such as methenyltetrahydrofolate, in
animal and plant cryptochromes is still under debate.
Arabidopsis cryptochrome 1 (CRY1) has been intensively
studied with regard to function and interaction of the
protein in vivo and in vitro. However, little is known about
the pathway from light absorption to signal transduction on
the molecular level. We investigated the full-length CRY1
protein by Fourier transform infrared (FTIR) and UV/vis
difference spectroscopy. Starting from the fully oxidized
state of the chromophore FAD, a neutral flavoprotein radical
is formed upon illumination in the absence of any exogenous
electron donor. A preliminary assignment of the chromophore
bands is presented. The FTIR difference spectrum reveals
only moderate changes in secondary structure of the
apoprotein in response to the photoreduction of the
chromophore. Deprotonation of an aspartic or glutamic acid,
probably D396, accompanies radical formation, as is deduced
from the negative band at 1734 cm(-)(1) in D(2)O. The main
positive band at 1524 cm(-)(1) in the FTIR spectrum shows a
strong shift to lower frequencies as compared to other
flavoproteins. Together with the unusual blue-shift of the
absorption in the visible range to 595 nm, this clearly
distinguishes the radical form of CRY1 from those of
structurally highly homologous DNA photolyases. As a
consequence, the direct comparison of cryptochrome to
photolyase in terms of photoreactivity and mechanism has to
be made with caution.},
keywords = {Arabidopsis Proteins: chemistry / Cryptochromes /
Crystallography, X-Ray / Deoxyribodipyrimidine Photo-Lyase:
chemistry / Deoxyribodipyrimidine Photo-Lyase: metabolism /
Escherichia coli: metabolism / Flavins: metabolism /
Flavoproteins: chemistry / Flavoproteins: metabolism / Light
/ Models, Molecular / Signal Transduction: physiology /
Spectrophotometry, Ultraviolet / Spectroscopy, Fourier
Transform Infrared: methods / Structural Homology, Protein /
Arabidopsis Proteins (NLM Chemicals) / Cryptochromes (NLM
Chemicals) / Flavins (NLM Chemicals) / Flavoproteins (NLM
Chemicals) / Deoxyribodipyrimidine Photo-Lyase (NLM
Chemicals) / J (WoSType)},
cin = {IBI-2},
ddc = {570},
cid = {I:(DE-Juel1)VDB58},
pnm = {Funktion und Dysfunktion des Nervensystems},
pid = {G:(DE-Juel1)FUEK409},
shelfmark = {Biochemistry $\&$ Molecular Biology},
typ = {PUB:(DE-HGF)16},
pubmed = {pmid:16489739},
UT = {WOS:000235792300002},
doi = {10.1021/bi051964b},
url = {https://juser.fz-juelich.de/record/51338},
}