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@ARTICLE{Braun:820059,
author = {Braun, Tatjana and Vos, Matthijn R. and Kalisman, Nir and
Sherman, Nicholas E. and Rachel, Reinhard and Wirth,
Reinhard and Schröder, Gunnar and Egelman, Edward H.},
title = {{A}rchaeal flagellin combines a bacterial type {IV} pilin
domain with an {I}g-like domain},
journal = {Proceedings of the National Academy of Sciences of the
United States of America},
volume = {113},
number = {37},
issn = {1091-6490},
address = {Washington, DC},
publisher = {National Acad. of Sciences},
reportid = {FZJ-2016-05616},
pages = {10352 - 10357},
year = {2016},
abstract = {The bacterial flagellar apparatus, which involves ∼40
different proteins, has been a model system for
understanding motility and chemotaxis. The bacterial
flagellar filament, largely composed of a single protein,
flagellin, has been a model for understanding protein
assembly. This system has no homology to the eukaryotic
flagellum, in which the filament alone, composed of a
microtubule-based axoneme, contains more than 400 different
proteins. The archaeal flagellar system is simpler still, in
some cases having ∼13 different proteins with a single
flagellar filament protein. The archaeal flagellar system
has no homology to the bacterial one and must have arisen by
convergent evolution. However, it has been understood that
the N-terminal domain of the archaeal flagellin is a homolog
of the N-terminal domain of bacterial type IV pilin, showing
once again how proteins can be repurposed in evolution for
different functions. Using cryo-EM, we have been able to
generate a nearly complete atomic model for a flagellar-like
filament of the archaeon Ignicoccus hospitalis from a
reconstruction at ∼4-Å resolution. We can now show that
the archaeal flagellar filament contains a β-sandwich,
previously seen in the FlaF protein that forms the anchor
for the archaeal flagellar filament. In contrast to the
bacterial flagellar filament, where the outer globular
domains make no contact with each other and are not
necessary for either assembly or motility, the archaeal
flagellin outer domains make extensive contacts with each
other that largely determine the interesting mechanical
properties of these filaments, allowing these filaments to
flex.},
cin = {ICS-6},
ddc = {000},
cid = {I:(DE-Juel1)ICS-6-20110106},
pnm = {551 - Functional Macromolecules and Complexes (POF3-551)},
pid = {G:(DE-HGF)POF3-551},
typ = {PUB:(DE-HGF)16},
UT = {WOS:000383092000046},
pubmed = {pmid:27578865},
doi = {10.1073/pnas.1607756113},
url = {https://juser.fz-juelich.de/record/820059},
}