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@ARTICLE{Fischer:827054,
author = {Fischer, Niels and Neumann, Piotr and Bock, Lars V. and
Maracci, Cristina and Wang, Zhe and Paleskava, Alena and
Konevega, Andrey L. and Schröder, Gunnar and Helmut,
Grubmüller and Ficner, Ralf and Rodnina, Marina V. and
Stark, Holger},
title = {{T}he pathway to {GTP}ase activation of elongation factor
{S}el{B} on the ribosome},
journal = {Nature},
volume = {540},
issn = {0028-0836},
address = {London [u.a.]},
publisher = {Nature Publ. Group},
reportid = {FZJ-2017-01260},
pages = {80–85},
year = {2016},
abstract = {In all domains of life, selenocysteine (Sec) is delivered
to the ribosome by selenocysteine-specific tRNA (tRNASec)
with the help of a specialized translation factor, SelB in
bacteria. Sec-tRNASec recodes a UGA stop codon next to a
downstream mRNA stem–loop. Here we present the structures
of six intermediates on the pathway of UGA recoding in
Escherichia coli by single-particle cryo-electron
microscopy. The structures explain the specificity of
Sec-tRNASec binding by SelB and show large-scale
rearrangements of Sec-tRNASec. Upon initial binding of
SelB–Sec-tRNASec to the ribosome and codon reading, the
30S subunit adopts an open conformation with Sec-tRNASec
covering the sarcin–ricin loop (SRL) on the 50S subunit.
Subsequent codon recognition results in a local closure of
the decoding site, which moves Sec-tRNASec away from the SRL
and triggers a global closure of the 30S subunit shoulder
domain. As a consequence, SelB docks on the SRL, activating
the GTPase of SelB. These results reveal how codon
recognition triggers GTPase activation in translational
GTPases.},
cin = {ICS-6},
ddc = {070},
cid = {I:(DE-Juel1)ICS-6-20110106},
pnm = {551 - Functional Macromolecules and Complexes (POF3-551)},
pid = {G:(DE-HGF)POF3-551},
typ = {PUB:(DE-HGF)16},
UT = {WOS:000388916600051},
pubmed = {pmid:27842381},
doi = {10.1038/nature20560},
url = {https://juser.fz-juelich.de/record/827054},
}