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@ARTICLE{Thorpe:8364,
author = {Thorpe, M.R. and Furch, A.C.U. and Minchin, P.E.H. and
Föller, J. and Van Bel, A.J.E. and Hafke, J.B.},
title = {{R}apid cooling triggers forisome dispersion just before
phloem transport stops},
journal = {Plant, cell $\&$ environment},
volume = {33},
issn = {0140-7791},
address = {Oxford [u.a.]},
publisher = {Wiley-Blackwell},
reportid = {PreJuSER-8364},
pages = {259 - 271},
year = {2010},
note = {We were supported by the Deutsche Forschungsgemeinschaft in
the frame of the Schwerpunktprogramm 1108 (BE 1925/8-2, 8-3,
15-1) and by the Marsden Fund of New Zealand (Contract
Number UOW003). We acknowledge Gerhard Roeb, Siegfried
Jahnke and Marco Dautzenberg for discussions and technical
assistance in Julich, and Werner Uhmann, Thomas Wagner,
Andreas Reh and Stefan Balser in Giessen for expert workshop
assistance in Giessen.},
abstract = {Phloem transport stops transiently within dicot stems that
are cooled rapidly, but the cause remains unknown. Now it is
known that (1) rapid cooling depolarizes cell membranes
giving a transient increase in cytoplasmic Ca(2+), and (2) a
rise of free calcium triggers dispersion of forisomes, which
then occlude sieve elements (SEs) of fabacean plants.
Therefore, we compared the effects of rapid chilling on SE
electrophysiology, phloem transport and forisomes in Vicia
faba. Forisomes dispersed after rapid cooling with a delay
that was longer for slower cooling rates. Phloem transport
stopped about 20 s after forisome dispersion, and then
transport resumed and forisomes re-condensed within similar
time frames. Transport interruption and forisome dispersion
showed parallel behaviour--a cooling rate-dependent
response, transience and desensitization. Chilling induced
both a fast and a slow depolarization of SE membranes, the
electrical signature suggesting strongly that the cause of
forisome dispersion was the transient promotion of SE free
calcium. This apparent block of SEs by dispersed forisomes
may be assisted by other Ca(2+)-dependent sealing proteins
that are present in all dicots.},
keywords = {Calcium: metabolism / Carbon Isotopes: analysis / Cold
Temperature / Electrophysiology / Fluorescent Dyes /
Membrane Potentials / Microelectrodes / Microscopy, Confocal
/ Phloem: physiology / Vicia faba: physiology / Carbon
Isotopes (NLM Chemicals) / Fluorescent Dyes (NLM Chemicals)
/ Calcium (NLM Chemicals) / J (WoSType)},
cin = {ICG-3},
ddc = {570},
cid = {I:(DE-Juel1)ICG-3-20090406},
pnm = {Terrestrische Umwelt},
pid = {G:(DE-Juel1)FUEK407},
shelfmark = {Plant Sciences},
typ = {PUB:(DE-HGF)16},
pubmed = {pmid:19930129},
UT = {WOS:000273551100010},
doi = {10.1111/j.1365-3040.2009.02079.x},
url = {https://juser.fz-juelich.de/record/8364},
}