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@ARTICLE{Theers:856922,
      author       = {Theers, Mario and Westphal, Elmar and Qi, Kai and Winkler,
                      Roland G. and Gompper, Gerhard},
      title        = {{C}lustering of microswimmers: interplay of shape and
                      hydrodynamics},
      journal      = {Soft matter},
      volume       = {14},
      number       = {42},
      issn         = {1744-6848},
      address      = {London},
      publisher    = {Royal Soc. of Chemistry},
      reportid     = {FZJ-2018-06249},
      pages        = {8590 - 8603},
      year         = {2018},
      abstract     = {The spatiotemporal dynamics in systems of active
                      self-propelled particles is controlled by the propulsion
                      mechanism in combination with various direct interactions,
                      such as steric repulsion and hydrodynamics. These direct
                      interactions are typically anisotropic, and come in
                      different “flavors”, such as spherical and elongated
                      particle shapes, pusher and puller flow fields, etc. The
                      combination of the various aspects is expected to lead to
                      new emergent behavior. However, it is a priori not evident
                      whether shape and hydrodynamics act synergistically or
                      antagonistically to generate motility-induced clustering
                      (MIC) and phase separation (MIPS). We employ a model of
                      prolate spheroidal microswimmers—called squirmers—in
                      quasi-two-dimensional confinement to address this issue by
                      mesoscale hydrodynamic simulations. For comparison,
                      non-hydrodynamic active Brownian particles (ABPs) are
                      considered to elucidate the contribution of hydrodynamic
                      interactions. For spherical particles, the comparison
                      between ABPs and hydrodynamic-squirmer ensembles reveals a
                      suppression of MIPS due to hydrodynamic interactions. Yet,
                      our analysis shows that dynamic clusters exist, with a broad
                      size distribution. The fundamental difference between ABPs
                      and squirmers is attributed to an increased reorientation of
                      squirmers by hydrodynamic torques during their collisions.
                      In contrast, for elongated squirmers, hydrodynamics
                      interactions enhance MIPS. The transition to a
                      phase-separated state strongly depends on the nature of the
                      swimmer's flow field—with an increased tendency toward
                      MIPS for pullers, and a reduced tendency for pushers. Thus,
                      hydrodynamic interactions show opposing effects on MIPS for
                      spherical and elongated microswimmers, and details of the
                      propulsion mechanism of biological microswimmers may be very
                      important to determine their collective behavior.},
      cin          = {IAS-2 / JARA-HPC},
      ddc          = {530},
      cid          = {I:(DE-Juel1)IAS-2-20090406 / $I:(DE-82)080012_20140620$},
      pnm          = {553 - Physical Basis of Diseases (POF3-553) / Collective
                      Dynamics of Microswimmers $(jias21_20171101)$},
      pid          = {G:(DE-HGF)POF3-553 / $G:(DE-Juel1)jias21_20171101$},
      typ          = {PUB:(DE-HGF)16},
      pubmed       = {pmid:30339172},
      UT           = {WOS:000448948800012},
      doi          = {10.1039/C8SM01390J},
      url          = {https://juser.fz-juelich.de/record/856922},
}