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@ARTICLE{Verma:865808,
author = {Verma, Neha and Dollinger, Peter and Kovacic, Filip and
Jaeger, Karl-Erich and Gohlke, Holger},
title = {{T}he {M}embrane-{I}ntegrated {S}teric {C}haperone {L}if
{F}acilitates {A}ctive {S}ite {O}pening of {P}seudomonas
aeruginosa {L}ipase {A}},
journal = {Journal of computational chemistry},
volume = {4},
number = {6},
issn = {0192-8651},
address = {New York, NY [u.a.]},
publisher = {Wiley},
reportid = {FZJ-2019-05106},
pages = {500-512},
year = {2020},
abstract = {Lipases are essential and widely used biocatalysts. Hence,
the production of lipases requires a detailed understanding
of the molecular mechanism of its folding and secretion.
Lipase A from Pseudomonas aeruginosa, PaLipA, constitutes a
prominent example that has additional relevance because of
its role as a virulence factor in many diseases. PaLipA
requires the assistance of a membrane‐integrated steric
chaperone, the lipase‐specific foldase Lif, to achieve its
enzymatically active state. However, the molecular mechanism
of how Lif activates its cognate lipase has remained
elusive. Here, we show by molecular dynamics simulations at
the atomistic level and potential of mean force computations
that Lif catalyzes the activation process of PaLipA by
structurally stabilizing an intermediate PaLipA
conformation, particularly a β‐sheet in the region of
residues 17–30, such that the opening of PaLipA's lid
domain is facilitated. This opening allows substrate access
to PaLipA's catalytic site. A surprising and so far not
fully understood aspect of our study is that the open state
of PaLipA is unstable compared to the closed one according
to our computational and in vitro biochemical results. We
thus speculate that further interactions of PaLipA with the
Xcp secretion machinery and/or components of the
extracellular matrix contribute to the remaining activity of
secreted PaLipA.},
cin = {NIC / JSC / ICS-6 / IMET},
ddc = {540},
cid = {I:(DE-Juel1)NIC-20090406 / I:(DE-Juel1)JSC-20090406 /
I:(DE-Juel1)ICS-6-20110106 / I:(DE-Juel1)IMET-20090612},
pnm = {511 - Computational Science and Mathematical Methods
(POF3-511) / Forschergruppe Gohlke $(hkf7_20170501)$ /
Analysis of the conformational changes during activation of
lipase A by its foldase $(hdd16_20171101)$ / Conformational
dynamics of the unbound lipase-specific foldase Lif
$(hdd16_20161101)$},
pid = {G:(DE-HGF)POF3-511 / $G:(DE-Juel1)hkf7_20170501$ /
$G:(DE-Juel1)hdd16_20171101$ / $G:(DE-Juel1)hdd16_20161101$},
typ = {PUB:(DE-HGF)16},
pubmed = {pmid:31618459},
UT = {WOS:000490396800001},
doi = {10.1002/jcc.26085},
url = {https://juser.fz-juelich.de/record/865808},
}