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@ARTICLE{Bruce:866099,
author = {Bruce, Neil J. and Narzi, Daniele and Trpevski, Daniel and
van Keulen, Siri C. and Nair, Anu G. and Röthlisberger,
Ursula and Wade, Rebecca C. and Carloni, Paolo and Hellgren
Kotaleski, Jeanette},
title = {{R}egulation of adenylyl cyclase 5 in striatal neurons
confers the ability to detect coincident neuromodulatory
signals},
journal = {PLoS Computational Biology},
volume = {15},
number = {10},
issn = {1553-7358},
address = {San Francisco, Calif.},
publisher = {Public Library of Science},
reportid = {FZJ-2019-05318},
pages = {e1007382 -},
year = {2019},
abstract = {Long-term potentiation and depression of synaptic activity
in response to stimuli is a key factor in reinforcement
learning. Strengthening of the corticostriatal synapses
depends on the second messenger cAMP, whose synthesis is
catalysed by the enzyme adenylyl cyclase 5 (AC5), which is
itself regulated by the stimulatory Gαolf and inhibitory
Gαi proteins. AC isoforms have been suggested to act as
coincidence detectors, promoting cellular responses only
when convergent regulatory signals occur close in time.
However, the mechanism for this is currently unclear, and
seems to lie in their diverse regulation patterns. Despite
attempts to isolate the ternary complex, it is not known if
Gαolf and Gαi can bind to AC5 simultaneously, nor what
activity the complex would have. Using protein
structure-based molecular dynamics simulations, we show that
this complex is stable and inactive. These simulations,
along with Brownian dynamics simulations to estimate protein
association rates constants, constrain a kinetic model that
shows that the presence of this ternary inactive complex is
crucial for AC5’s ability to detect coincident signals,
producing a synergistic increase in cAMP. These results
reveal some of the prerequisites for corticostriatal
synaptic plasticity, and explain recent experimental data on
cAMP concentrations following receptor activation. Moreover,
they provide insights into the regulatory mechanisms that
control signal processing by different AC isoforms.},
cin = {IAS-5 / INM-9},
ddc = {610},
cid = {I:(DE-Juel1)IAS-5-20120330 / I:(DE-Juel1)INM-9-20140121},
pnm = {574 - Theory, modelling and simulation (POF3-574)},
pid = {G:(DE-HGF)POF3-574},
typ = {PUB:(DE-HGF)16},
pubmed = {pmid:31665146},
UT = {WOS:000500776600040},
doi = {10.1371/journal.pcbi.1007382},
url = {https://juser.fz-juelich.de/record/866099},
}