000008698 001__ 8698 000008698 005__ 20200402205809.0 000008698 0247_ $$2pmid$$apmid:20090755 000008698 0247_ $$2pmc$$apmc:PMC2834796 000008698 0247_ $$2DOI$$a10.1038/nature08701 000008698 0247_ $$2WOS$$aWOS:000273748100049 000008698 0247_ $$2altmetric$$aaltmetric:519603 000008698 037__ $$aPreJuSER-8698 000008698 041__ $$aeng 000008698 082__ $$a070 000008698 084__ $$2WoS$$aMultidisciplinary Sciences 000008698 1001_ $$0P:(DE-HGF)0$$aZhang, J.$$b0 000008698 245__ $$aMechanism of folding chamber closure in a group II chaperonin 000008698 260__ $$aLondon [u.a.]$$bNature Publising Group$$c2010 000008698 300__ $$a379 - 383 000008698 3367_ $$0PUB:(DE-HGF)16$$2PUB:(DE-HGF)$$aJournal Article 000008698 3367_ $$2DataCite$$aOutput Types/Journal article 000008698 3367_ $$00$$2EndNote$$aJournal Article 000008698 3367_ $$2BibTeX$$aARTICLE 000008698 3367_ $$2ORCID$$aJOURNAL_ARTICLE 000008698 3367_ $$2DRIVER$$aarticle 000008698 440_0 $$04484$$aNature$$v463$$x0028-0836 000008698 500__ $$aWe acknowledge the support of grants from the National Institutes of Health through the Nanomedicine Development Center Roadmap Initiative, Biomedical Technology Research Center for Structural Biology in National Center for Research Resources, Nanobiology Training Fellowship administered by the Keck Center of the Gulf Coast Consortia and the National Science Foundation. 000008698 520__ $$aGroup II chaperonins are essential mediators of cellular protein folding in eukaryotes and archaea. These oligomeric protein machines, approximately 1 megadalton, consist of two back-to-back rings encompassing a central cavity that accommodates polypeptide substrates. Chaperonin-mediated protein folding is critically dependent on the closure of a built-in lid, which is triggered by ATP hydrolysis. The structural rearrangements and molecular events leading to lid closure are still unknown. Here we report four single particle cryo-electron microscopy (cryo-EM) structures of Mm-cpn, an archaeal group II chaperonin, in the nucleotide-free (open) and nucleotide-induced (closed) states. The 4.3 A resolution of the closed conformation allowed building of the first ever atomic model directly from the single particle cryo-EM density map, in which we were able to visualize the nucleotide and more than 70% of the side chains. The model of the open conformation was obtained by using the deformable elastic network modelling with the 8 A resolution open-state cryo-EM density restraints. Together, the open and closed structures show how local conformational changes triggered by ATP hydrolysis lead to an alteration of intersubunit contacts within and across the rings, ultimately causing a rocking motion that closes the ring. Our analyses show that there is an intricate and unforeseen set of interactions controlling allosteric communication and inter-ring signalling, driving the conformational cycle of group II chaperonins. Beyond this, we anticipate that our methodology of combining single particle cryo-EM and computational modelling will become a powerful tool in the determination of atomic details involved in the dynamic processes of macromolecular machines in solution. 000008698 536__ $$0G:(DE-Juel1)FUEK409$$2G:(DE-HGF)$$aFunktion und Dysfunktion des Nervensystems$$cP33$$x0 000008698 536__ $$0G:(DE-Juel1)FUEK505$$aBioSoft: Makromolekulare Systeme und biologische Informationsverarbeitung$$cP45$$x1 000008698 588__ $$aDataset connected to Web of Science, Pubmed 000008698 650_2 $$2MeSH$$aAdenosine Triphosphate: chemistry 000008698 650_2 $$2MeSH$$aAdenosine Triphosphate: metabolism 000008698 650_2 $$2MeSH$$aAdenosine Triphosphate: pharmacology 000008698 650_2 $$2MeSH$$aAllosteric Regulation 000008698 650_2 $$2MeSH$$aBinding Sites 000008698 650_2 $$2MeSH$$aCryoelectron Microscopy 000008698 650_2 $$2MeSH$$aGroup II Chaperonins: chemistry 000008698 650_2 $$2MeSH$$aGroup II Chaperonins: metabolism 000008698 650_2 $$2MeSH$$aGroup II Chaperonins: ultrastructure 000008698 650_2 $$2MeSH$$aHydrolysis: drug effects 000008698 650_2 $$2MeSH$$aMethanococcus: chemistry 000008698 650_2 $$2MeSH$$aModels, Molecular 000008698 650_2 $$2MeSH$$aProtein Binding 000008698 650_2 $$2MeSH$$aProtein Conformation: drug effects 000008698 650_2 $$2MeSH$$aProtein Folding 000008698 650_2 $$2MeSH$$aProtein Subunits: chemistry 000008698 650_2 $$2MeSH$$aProtein Subunits: metabolism 000008698 650_2 $$2MeSH$$aStructure-Activity Relationship 000008698 650_7 $$00$$2NLM Chemicals$$aProtein Subunits 000008698 650_7 $$056-65-5$$2NLM Chemicals$$aAdenosine Triphosphate 000008698 650_7 $$0EC 3.6.1.-$$2NLM Chemicals$$aGroup II Chaperonins 000008698 650_7 $$2WoSType$$aJ 000008698 7001_ $$0P:(DE-HGF)0$$aBaker, M.L.$$b1 000008698 7001_ $$0P:(DE-Juel1)132018$$aSchröder, G.F.$$b2$$uFZJ 000008698 7001_ $$0P:(DE-HGF)0$$aDouglas, N.R.$$b3 000008698 7001_ $$0P:(DE-HGF)0$$aReissmann, S.$$b4 000008698 7001_ $$0P:(DE-HGF)0$$aJakana, J.$$b5 000008698 7001_ $$0P:(DE-HGF)0$$aDougherty, M.$$b6 000008698 7001_ $$0P:(DE-HGF)0$$aFu, C.J.$$b7 000008698 7001_ $$0P:(DE-HGF)0$$aLevitt, M.$$b8 000008698 7001_ $$0P:(DE-HGF)0$$aLudtke, S.J.$$b9 000008698 7001_ $$0P:(DE-HGF)0$$aFrydman, J.$$b10 000008698 7001_ $$0P:(DE-HGF)0$$aChiu, W.$$b11 000008698 773__ $$0PERI:(DE-600)1413423-8$$a10.1038/nature08701$$gVol. 463, p. 379 - 383$$p379 - 383$$q463<379 - 383$$tNature <London>$$v463$$x0028-0836$$y2010 000008698 8567_ $$2Pubmed Central$$uhttp://www.ncbi.nlm.nih.gov/pmc/articles/PMC2834796 000008698 909CO $$ooai:juser.fz-juelich.de:8698$$pVDB 000008698 9131_ $$0G:(DE-Juel1)FUEK409$$bGesundheit$$kP33$$lFunktion und Dysfunktion des Nervensystems$$vFunktion und Dysfunktion des Nervensystems$$x0 000008698 9131_ $$0G:(DE-Juel1)FUEK505$$bSchlüsseltechnologien$$kP45$$lBiologische Informationsverarbeitung$$vBioSoft: Makromolekulare Systeme und biologische Informationsverarbeitung$$x1 000008698 9132_ $$0G:(DE-HGF)POF3-551$$1G:(DE-HGF)POF3-550$$2G:(DE-HGF)POF3-500$$aDE-HGF$$bKey Technologies$$lBioSoft Fundamentals for future Technologies in the fields of Soft Matter and Life Sciences$$vFunctional Macromolecules and Complexes$$x0 000008698 9141_ $$y2010 000008698 915__ $$0StatID:(DE-HGF)0010$$aJCR/ISI refereed 000008698 9201_ $$0I:(DE-Juel1)VDB942$$d31.12.2010$$gISB$$kISB-3$$lStrukturbiochemie$$x0 000008698 970__ $$aVDB:(DE-Juel1)117765 000008698 980__ $$aVDB 000008698 980__ $$aConvertedRecord 000008698 980__ $$ajournal 000008698 980__ $$aI:(DE-Juel1)ICS-6-20110106 000008698 980__ $$aUNRESTRICTED 000008698 981__ $$aI:(DE-Juel1)IBI-7-20200312 000008698 981__ $$aI:(DE-Juel1)ICS-6-20110106