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@ARTICLE{Wolf:873824,
author = {Wolf, Natalie and Bussmann, Michael and Koch-Koerfges,
Abigail and Katcharava, Nino and Schulte, Julia and Polen,
Tino and Hartl, Johannes and Vorholt, Julia A. and Baumgart,
Meike and Bott, Michael},
title = {{M}olecular {B}asis of {G}rowth {I}nhibition by {A}cetate
of an {A}denylate {C}yclase-{D}eficient {M}utant of
{C}orynebacterium glutamicum},
journal = {Frontiers in microbiology},
volume = {11},
issn = {1664-302X},
address = {Lausanne},
publisher = {Frontiers Media},
reportid = {FZJ-2020-01029},
pages = {87},
year = {2020},
note = {Biotechnologie 1},
abstract = {In Corynebacterium glutamicum, cyclic adenosine
monophosphate (cAMP) serves as an effector of the global
transcriptional regulator GlxR. Synthesis of cAMP is
catalyzed by the membrane-bound adenylate cyclase CyaB. In
this study, we investigated the consequences of decreased
intracellular cAMP levels in a ΔcyaB mutant. While no
growth defect of the ΔcyaB strain was observed on glucose,
fructose, sucrose, or gluconate alone, the addition of
acetate to these growth media resulted in a severe growth
inhibition, which could be reversed by plasmid-based cyaB
expression or by supplementation of the medium with cAMP.
The effect was concentration- and pH-dependent, suggesting a
link to the uncoupling activity of acetate. In agreement,
the ΔcyaB mutant had an increased sensitivity to the
protonophore carbonyl cyanide m-chlorophenyl hydrazone
(CCCP). The increased uncoupler sensitivity correlated with
a lowered membrane potential of acetate-grown ΔcyaB cells
compared to wild-type cells. A reduced membrane potential
affects major cellular processes, such as ATP synthesis by
F1FO-ATP synthase and numerous transport processes. The
impaired membrane potential of the ΔcyaB mutant could be
due to a decreased expression of the cytochrome bc1-aa3
supercomplex, which is the major contributor of
proton-motive force in C. glutamicum. Expression of the
supercomplex genes was previously reported to be activated
by GlxR-cAMP. A suppressor mutant of the ΔcyaB strain with
improved growth on acetate was isolated, which carried a
single mutation in the genome leading to an Ala131Thr
exchange in GlxR. Introduction of this point mutation into
the original ΔcyaB mutant restored the growth defect on
acetate. This supported the importance of GlxR for the
phenotype of the ΔcyaB mutant and, more generally, of the
cAMP-GlxR system for the control of energy metabolism in C.
glutamicum.Introduction},
cin = {IBG-1},
ddc = {570},
cid = {I:(DE-Juel1)IBG-1-20101118},
pnm = {581 - Biotechnology (POF3-581)},
pid = {G:(DE-HGF)POF3-581},
typ = {PUB:(DE-HGF)16},
pubmed = {pmid:32117117},
UT = {WOS:000517630100001},
doi = {10.3389/fmicb.2020.00087},
url = {https://juser.fz-juelich.de/record/873824},
}