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@ARTICLE{DeBauw:878667,
author = {De Bauw, Pieterjan and Mai, Trung Hieu and Schnepf, Andrea
and Merckx, Roel and Smolders, Erik and Vanderborght, Jan},
title = {{A} functional–structural model of upland rice root
systems reveals the importance of laterals and growing root
tips for phosphate uptake from wet and dry soils},
journal = {Annals of botany},
volume = {126},
number = {4},
issn = {1095-8290},
address = {Oxford},
publisher = {Oxford University Press},
reportid = {FZJ-2020-02988},
pages = {789–806},
year = {2020},
abstract = {Background and AimsUpland rice is often grown where water
and phosphorus (P) are limited. To better understand the
interaction between water and P availability,
functional–structural models that mechanistically
represent small-scale nutrient gradients and water dynamics
in the rhizosphere are needed.MethodsRice was grown in large
columns using a P-deficient soil at three P supplies in the
topsoil (deficient, sub-optimal and non-limiting) in
combination with two water regimes (field capacity vs.
drying periods). Root system characteristics, such as nodal
root number, lateral types, interbranch distance, root
diameters and the distribution of biomass with depth, as
well as water and P uptake, were measured. Based on the
observed root data, 3-D root systems were reconstructed by
calibrating the structural architecure model CRootBox for
each scenario. Water flow and P transport in the soil to
each of the individual root segments of the generated 3-D
root architectures were simulated using a multiscale flow
and transport model. Total water and P uptake were then
computed by adding up the uptake by all the root
segments.Key ResultsMeasurements showed that root
architecture was significantly affected by the treatments.
The moist, high P scenario had 2.8 times the root mass,
double the number of nodal roots and more S-type laterals
than the dry, low P scenario. Likewise, measured plant P
uptake increased >3-fold by increasing P and water supply.
However, drying periods reduced P uptake at high but not at
low P supply. Simulation results adequately predicted P
uptake in all scenarios when the Michaelis–Menten constant
(Km) was corrected for diffusion limitation. They showed
that the key drivers for P uptake are the different types of
laterals (i.e. S- and L-type) and growing root tips. The
L-type laterals become more important for overall water and
P uptake than the S-type laterals in the dry scenarios. This
is true across all the P treatments, but the effect is more
pronounced as the P availability decreases.ConclusionsThis
functional–structural model can predict the function of
specific rice roots in terms of P and water uptake under
different P and water supplies, when the structure of the
root system is known. A future challenge is to predict how
the structure root systems responds to nutrient and water
availability.},
cin = {IBG-3},
ddc = {580},
cid = {I:(DE-Juel1)IBG-3-20101118},
pnm = {255 - Terrestrial Systems: From Observation to Prediction
(POF3-255)},
pid = {G:(DE-HGF)POF3-255},
typ = {PUB:(DE-HGF)16},
pubmed = {pmid:32597468},
UT = {WOS:000591847300020},
doi = {10.1093/aob/mcaa120},
url = {https://juser.fz-juelich.de/record/878667},
}