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@ARTICLE{Twizerimana:884114,
author = {Twizerimana, Augustin Penda and Scheck, Rachel and Becker,
Daniel and Zhang, Zeli and Wammers, Marianne and Avelar,
Leandro and Pflieger, Marc and Häussinger, Dieter and Kurz,
Thomas and Gohlke, Holger and Münk, Carsten},
title = {{C}ell type-dependent escape of capsid inhibitors by simian
immunodeficiency virus {SIV}cpz},
journal = {Journal of virology},
volume = {94},
number = {23},
issn = {1098-5514},
address = {Baltimore, Md.},
publisher = {Soc.},
reportid = {FZJ-2020-03101},
pages = {e01338-20},
year = {2020},
abstract = {Pandemic human immunodeficiency virus type 1 (HIV-1) is the
result of the zoonotic transmission of simian
immunodeficiency virus (SIV) from the chimpanzee subspecies
Pan troglodytes troglodytes (SIVcpzPtt). The related
subspecies Pan troglodytes schweinfurthii is the host of a
similar virus, SIVcpzPts, which did not spread to humans. We
tested these viruses with small-molecule capsid inhibitors
(PF57, PF74, and GS-CA1) that interact with a binding groove
in the capsid that is also used by CPSF6. While HIV-1 was
sensitive to capsid inhibitors in cell lines, human
macrophages, and peripheral blood mononuclear cells (PBMCs),
SIVcpzPtt was resistant in rhesus FRhL-2 cells and human
PBMCs but was sensitive to PF74 in human HOS and HeLa cells.
SIVcpzPts was insensitive to PF74 in FRhL-2 cells, HeLa
cells, PBMCs, and macrophages but was inhibited by PF74 in
HOS cells. A truncated version of CPSF6 (CPSF6-358)
inhibited SIVcpzPtt and HIV-1, while in contrast, SIVcpzPts
was resistant to CPSF6-358. Homology modeling of HIV-1,
SIVcpzPtt, and SIVcpzPts capsids and binding energy
estimates suggest that these three viruses bind similarly to
the host proteins cyclophilin A (CYPA) and CPSF6 as well as
the capsid inhibitor PF74. Cyclosporine treatment, mutation
of the CYPA-binding loop in the capsid, or CYPA knockout
eliminated the resistance of SIVcpzPts to PF74 in HeLa
cells. These experiments revealed that the antiviral
capacity of PF74 is controlled by CYPA in a virus- and cell
type-specific manner. Our data indicate that SIVcpz viruses
can use infection pathways that escape the antiviral
activity of PF74. We further suggest that the antiviral
activity of PF74 capsid inhibitors depends on cellular
cofactors.IMPORTANCE HIV-1 originated from SIVcpzPtt but not
from the related virus SIVcpzPts, and thus, it is important
to describe molecular infection by SIVcpzPts in human cells
to understand the zoonosis of SIVs. Pharmacological HIV-1
capsid inhibitors (e.g., PF74) bind a capsid groove that is
also a binding site for the cellular protein CPSF6.
SIVcpzPts was resistant to PF74 in HeLa cells but sensitive
in HOS cells, thus indicating cell line-specific resistance.
Both SIVcpz viruses showed resistance to PF74 in human
PBMCs. Modulating the presence of cyclophilin A or its
binding to capsid in HeLa cells overcame SIVcpzPts
resistance to PF74. These results indicate that early
cytoplasmic infection events of SIVcpzPts may differ between
cell types and affect, in an unknown manner, the antiviral
activity of capsid inhibitors. Thus, capsid inhibitors
depend on the activity or interaction of currently
uncharacterized cellular factors.},
cin = {JSC / NIC / IBI-7},
ddc = {610},
cid = {I:(DE-Juel1)JSC-20090406 / I:(DE-Juel1)NIC-20090406 /
I:(DE-Juel1)IBI-7-20200312},
pnm = {511 - Computational Science and Mathematical Methods
(POF3-511) / Forschergruppe Gohlke $(hkf7_20170501)$},
pid = {G:(DE-HGF)POF3-511 / $G:(DE-Juel1)hkf7_20170501$},
typ = {PUB:(DE-HGF)16},
pubmed = {32907979},
UT = {WOS:000595866900008},
doi = {10.1128/JVI.01338-20},
url = {https://juser.fz-juelich.de/record/884114},
}