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@PHDTHESIS{Kraxner:888609,
author = {Kraxner, Kim},
title = {{N}ovel insights into the transcriptional regulation of
cell division in {C}orynebacterium glutamicum},
volume = {241},
school = {Heinrich-Heine-Universität Düsseldorf},
type = {Dissertation},
address = {Jülich},
publisher = {Forschungszentrum Jülich GmbH Zentralbibliothek, Verlag},
reportid = {FZJ-2020-05065},
isbn = {978-3-95806-560-4},
series = {Schriften des Forschungszentrums Jülich Reihe
Schlüsseltechnologien / Key Technologies},
pages = {V, 83},
year = {2021},
note = {Biotechnologie 1; Dissertation, Heinrich-Heine-Universität
Düsseldorf, 2020},
abstract = {In the first part of this doctoral thesis the
transcriptional regulation of the odhI gene (cg1630)of
Corynebacterium glutamicum was analyzed. OdhI in its
unphoshorylated state functions asinhibitor of the
2-oxoglutarate dehydrogenase complex (ODHC) by binding to
the OdhAsubunit. Phosphorylation of OdhI by serine/threonine
protein kinases abolishes this effect.Inhibition of ODHC
activity by OdhI was shown to be crucial for overproduction
and secretionof L-glutamate, which is used as a flavour
enhancer. Since downstream of odhI two genespresumably
encoding transcriptional regulators (cg1631 and cg1633) are
located, it wasspeculated that these could be involved in
transcriptional regulation of odhI. However,transcriptome
analysis of deletion mutants lacking cg1631 or cg1633 and
DNA affinitychromatography with the odhI promoter did not
support this hypothesis. Furthermore, no otherpotential
transcriptional regulators of odhI could be identified.
Thus, there is currently noevidence for transcriptional
regulation of odhI.The second part of this thesis addresses
the regulation of cytokinesis in C. glutamicum. Incontrast
to e.g. Escherichia coli and Bacillus subtilis, knowledge
about regulators of cytokinesisin Actinobacteria is very
limited. In this study, the so far uncharacterized Cg1631
protein wasdiscovered to be a transcriptional regulator of
the ftsZ gene in C. glutamicum encoding the keyplayer of
bacterial cell division. Therefore, Cg1631 was named FtsR,
standing for FtsZregulator. Both deletion and overexpression
of ftsR caused growth defects and an altered cellmorphology,
emphasizing an important function of FtsR in cell division
or cell wall synthesis.The wild-type phenotype could be
restored by plasmid-based complementation. Chromatinaffinity
purification with subsequent next generation sequencing
(ChAP-Seq) identified a regionin the ftsZ promoter as a
major FtsR binding site, but revealed also additional
potential targetgenes. With the ChAP-Seq results a putative
DNA-binding motif could be identified for
FtsR.Transcriptional activation of ftsZ expression by FtsR
was underlined by DNA microarrayexperiments, electrophoretic
mobility shift assays (EMSAs), and reporter gene
studies.Analysis of strains expressing ftsZ under control of
the gluconate-inducible gntK promoterrevealed that the
phenotype of the ftsR mutant is not solely caused by
reduced ftsZexpression but involves additional factors. In
summary, FtsR was identified as the firsttranscriptional
regulator of ftsZ in C. glutamicum. Furthermore, since FtsR
and its DNA-bindingsite in the promoter region of ftsZ are
highly conserved in Actinobacteria, it can be assumedthat
this regulatory mechanism is also relevant for the control
of cell division in relatedActinobacteria. This makes FtsR a
promising target for the development of new
antimicrobialdrugs against pathogenic relatives of C.
glutamicum},
cin = {IBG-1},
cid = {I:(DE-Juel1)IBG-1-20101118},
pnm = {581 - Biotechnology (POF3-581)},
pid = {G:(DE-HGF)POF3-581},
typ = {PUB:(DE-HGF)3 / PUB:(DE-HGF)11},
urn = {urn:nbn:de:0001-2021100106},
url = {https://juser.fz-juelich.de/record/888609},
}