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@ARTICLE{Georgy:904312,
author = {Georgy, Jacqueline and Arlt, Yvonne and Moll, Jens M. and
Ouzin, Meryem and Weitz, Hendrik T. and Gremer, Lothar and
Willbold, Dieter and Grötzinger, Joachim and
Thives-Kurenbach, Felix and Scheller, Jürgen and Floss,
Doreen M.},
title = {{T}ryptophan ({W}) at position 37 of murine {IL}-12/{IL}-23
p40 is mandatory for binding to {IL}-12{R}β1 and subsequent
signal transduction},
journal = {The journal of biological chemistry},
volume = {297},
number = {5},
issn = {0021-9258},
address = {Bethesda, Md.},
publisher = {Soc.},
reportid = {FZJ-2021-05882},
pages = {101295 -},
year = {2021},
abstract = {Interleukin (IL)-12 and IL-23 are composite cytokines
consisting of p35/p40 and p19/p40, respectively, which
signal via the common IL-12 receptor β1 (IL-12Rβ1) and the
cytokine-specific receptors IL-12Rβ2 and IL-23R. Previous
data showed that the p40 component interacts with IL-12Rβ1,
whereas p19 and p35 subunits solely bind to IL-23R and
IL-12Rβ2, resulting in tetrameric signaling complexes. In
the absence of p19 and p35, p40 forms homodimers and may
induce signaling via IL-12Rβ1 homodimers. The critical
amino acids of p19 and p35 required for binding to IL-23R
and IL-12Rβ2 are known, and two regions of p40 critical for
binding to IL-12Rβ1 have recently been identified. In order
to characterize the involvement of the N-terminal region of
p40 in binding to IL-12Rβ1, we generated deletion variants
of the p40-p19 fusion cytokine. We found that an N-terminal
deletion variant missing amino acids M23 to P39 failed to
induce IL-23-dependent signaling and did not bind to
IL-12Rβ1, whereas binding to IL-23R was maintained. Amino
acid replacements showed that p40W37K largely abolished
IL-23-induced signal transduction and binding to IL-12Rβ1,
but not binding to IL-23R. Combining p40W37K with D36K and
T38K mutations eliminated the biological activity of IL-23.
Finally, homodimeric p40D36K/W37K/T38K did not interact with
IL-12Rβ1, indicating binding of homodimeric p40 to
IL-12Rβ1 is comparable to the interaction of IL-23/IL-12
and IL-12Rβ1. In summary, we have defined D36, W37, and T38
as hotspot amino acids for the interaction of IL-12/IL-23
p40 with IL-12Rβ1. Structural insights into
cytokine–cytokine receptor binding are important to
develop novel therapeutic strategies.},
cin = {IBI-7},
ddc = {540},
cid = {I:(DE-Juel1)IBI-7-20200312},
pnm = {5244 - Information Processing in Neuronal Networks
(POF4-524)},
pid = {G:(DE-HGF)POF4-5244},
typ = {PUB:(DE-HGF)16},
pubmed = {34637790},
UT = {WOS:000723057300007},
doi = {10.1016/j.jbc.2021.101295},
url = {https://juser.fz-juelich.de/record/904312},
}