000909777 001__ 909777 000909777 005__ 20240213111711.0 000909777 0247_ $$2doi$$a10.1038/s41467-022-32102-9 000909777 0247_ $$2Handle$$a2128/31858 000909777 0247_ $$2pmid$$a35977954 000909777 0247_ $$2WOS$$aWOS:001124833500001 000909777 037__ $$aFZJ-2022-03407 000909777 082__ $$a500 000909777 1001_ $$0P:(DE-HGF)0$$aSri-Ranjan, K.$$b0$$eCorresponding author 000909777 245__ $$aIntrinsic cell rheology drives junction maturation 000909777 260__ $$a[London]$$bNature Publishing Group UK$$c2022 000909777 3367_ $$2DRIVER$$aarticle 000909777 3367_ $$2DataCite$$aOutput Types/Journal article 000909777 3367_ $$0PUB:(DE-HGF)16$$2PUB:(DE-HGF)$$aJournal Article$$bjournal$$mjournal$$s1663655147_5639 000909777 3367_ $$2BibTeX$$aARTICLE 000909777 3367_ $$2ORCID$$aJOURNAL_ARTICLE 000909777 3367_ $$00$$2EndNote$$aJournal Article 000909777 520__ $$aA fundamental property of higher eukaryotes that underpins their evolutionary success is stable cell-cell cohesion. Yet, how intrinsic cell rheology and stiffness contributes to junction stabilization and maturation is poorly understood. We demonstrate that localized modulation of cell rheology governs the transition of a slack, undulated cell-cell contact (weak adhesion) to a mature, straight junction (optimal adhesion). Cell pairs confined on different geometries have heterogeneous elasticity maps and control their own intrinsic rheology co-ordinately. More compliant cell pairs grown on circles have slack contacts, while stiffer triangular cell pairs favour straight junctions with flanking contractile thin bundles. Counter-intuitively, straighter cell-cell contacts have reduced receptor density and less dynamic junctional actin, suggesting an unusual adaptive mechano-response to stabilize cell-cell adhesion. Our modelling informs that slack junctions arise from failure of circular cell pairs to increase their own intrinsic stiffness and resist the pressures from the neighbouring cell. The inability to form a straight junction can be reversed by increasing mechanical stress artificially on stiffer substrates. Our data inform on the minimal intrinsic rheology to generate a mature junction and provide a springboard towards understanding elements governing tissue-level mechanics. 000909777 536__ $$0G:(DE-HGF)POF4-333$$a333 - Integrative Biomedizin (POF4-333)$$cPOF4-333$$fPOF IV$$x0 000909777 536__ $$0G:(DE-HGF)POF4-315$$a315 - Bildgebung und Radioonkologie (POF4-315)$$cPOF4-315$$fPOF IV$$x1 000909777 536__ $$0G:(DE-HGF)POF4-5243$$a5243 - Information Processing in Distributed Systems (POF4-524)$$cPOF4-524$$fPOF IV$$x2 000909777 536__ $$0G:(GEPRIS)273723265$$aDFG project 273723265 - Mechanosensation und Mechanoreaktion in epidermalen Systemen $$c273723265$$x3 000909777 588__ $$aDataset connected to CrossRef, Journals: juser.fz-juelich.de 000909777 7001_ $$00000-0003-2942-0079$$aSanchez-Alonso, J. L.$$b1 000909777 7001_ $$0P:(DE-HGF)0$$aSwiatlowska, P.$$b2 000909777 7001_ $$0P:(DE-HGF)0$$aRothery, S.$$b3 000909777 7001_ $$00000-0002-5461-0344$$aNovak, P.$$b4 000909777 7001_ $$0P:(DE-Juel1)171121$$aGerlach, S.$$b5 000909777 7001_ $$0P:(DE-Juel1)188726$$aKoeninger, D.$$b6 000909777 7001_ $$0P:(DE-Juel1)128817$$aHoffmann, Bernd$$b7 000909777 7001_ $$0P:(DE-Juel1)128833$$aMerkel, R.$$b8 000909777 7001_ $$0P:(DE-HGF)0$$aStevens, M. M.$$b9 000909777 7001_ $$00000-0002-9077-7088$$aSun, S. X.$$b10 000909777 7001_ $$00000-0003-1148-9158$$aGorelik, J.$$b11 000909777 7001_ $$00000-0003-0546-7163$$aBraga, Vania M. 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