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@ARTICLE{Botte:825205,
author = {Botte, Mathieu and Zaccai, Nathan R. and Nijeholt, Jelger
Lycklama à. and Martin, Remy and Knoops, Kèvin and Papai,
Gabor and Zou, Juan and Deniaud, Aurélien and Karuppasamy,
Manikandan and Jiang, Qiyang and Roy, Abhishek Singha and
Schulten, Klaus and Schultz, Patrick and Rappsilber, Juri
and Zaccai, Giuseppe and Berger, Imre and Collinson, Ian and
Schaffitzel, Christiane},
title = {{A} central cavity within the holo-translocon suggests a
mechanism for membrane protein insertion},
journal = {Scientific reports},
volume = {6},
issn = {2045-2322},
address = {London},
publisher = {Nature Publishing Group},
reportid = {FZJ-2016-07676},
pages = {38399 -},
year = {2016},
abstract = {The conserved SecYEG protein-conducting channel and the
accessory proteins SecDF-YajC and YidC constitute the
bacterial holo-translocon (HTL), capable of
protein-secretion and membrane-protein insertion. By
employing an integrative approach combining small-angle
neutron scattering (SANS), low-resolution electron
microscopy and biophysical analyses we determined the
arrangement of the proteins and lipids within the
super-complex. The results guided the placement of X-ray
structures of individual HTL components and allowed the
proposal of a model of the functional translocon. Their
arrangement around a central lipid-containing pool conveys
an unexpected, but compelling mechanism for membrane-protein
insertion. The periplasmic domains of YidC and SecD are
poised at the protein-channel exit-site of SecY, presumably
to aid the emergence of translocating polypeptides. The SecY
lateral gate for membrane-insertion is adjacent to the
membrane ‘insertase’ YidC. Absolute-scale SANS employing
a novel contrast-match-point analysis revealed a dynamic
complex adopting open and compact configurations around an
adaptable central lipid-filled chamber, wherein polytopic
membrane-proteins could fold, sheltered from aggregation and
proteolysis.},
cin = {JCNS (München) ; Jülich Centre for Neutron Science JCNS
(München) ; JCNS-FRM-II},
ddc = {000},
cid = {I:(DE-Juel1)JCNS-FRM-II-20110218},
pnm = {6G15 - FRM II / MLZ (POF3-6G15) / 6G4 - Jülich Centre for
Neutron Research (JCNS) (POF3-623)},
pid = {G:(DE-HGF)POF3-6G15 / G:(DE-HGF)POF3-6G4},
experiment = {EXP:(DE-MLZ)KWS2-20140101},
typ = {PUB:(DE-HGF)16},
UT = {WOS:000389740800001},
pubmed = {pmid:27924919},
doi = {10.1038/srep38399},
url = {https://juser.fz-juelich.de/record/825205},
}